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The fossil record (especially the sister species, Pinguinus alfrednewtoni) and molecular evidence show that the three closely related genera diverged soon after their common ancestor, a bird probably similar to a stout Xantus's murrelet, had spread to the coasts of the Atlantic.

Apparently, by that time, the murres, or Atlantic guillemots, already had split from the other Atlantic alcids.

The Irish name for the great auk is 'falcóg mhór', meaning 'big seabird/auk'. This has led to an alternative English common name for the bird, "garefowl" or "gairfowl".

Although the etymology is debated, the generic name, "penguin", may be derived from the Welsh pen gwyn "white head", either because the birds lived in Newfoundland on White Head Island (Pen Gwyn in Welsh), or, because the great auk had such large white circles on its head.

Its favourite prey were fish, including Atlantic menhaden and capelin, and crustaceans.

Instead, the great auk was a powerful swimmer, a trait that it used in hunting.

Scientists soon began to realize that the great auk was disappearing and it became the beneficiary of many early environmental laws, but this proved ineffectual.

Its growing rarity increased interest from European museums and private collectors in obtaining skins and eggs of the bird.

Early European explorers to the Americas used the great auk as a convenient food source or as fishing bait, reducing its numbers.

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The bird's down was in high demand in Europe, a factor that largely eliminated the European populations by the mid-16th century.Razorbill-like birds were common in the Atlantic during the Pliocene, but the evolution of the little auk is sparsely documented.

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